Personal profile


Cand. Scient. (M.Sc., Biology-Chemistry), Odense University (February 1982).
Dr. Scient. (D.Sc.), Odense University (October 1988).

1982-1983: Adjunct, Sønderborg Statsskole. 1983-1987: Candidate and Senior Stipend, Institute of Biology, Odense University. 1987-1991: Assistant Professor, Institute of Biology, Odense University. 1991-2017: Associate Professor, Institute of Biology, Odense University (now University of Southern Denmark). 2018: Associate Professor emeritus, Institute of Biology, University of Southern Denmark. 1996-1998: Head of the Department (Institute Leader), Institute of Biology, Odense.

Visiting scientist: Department of Zoology, Helsinki University, Finland (1984, 1990); Department of Physiology, Max-Planck-Institute for Experimental Medicine, Göttingen, Germany (1986, 1988); Kristineberg Marine Research Station, Sweden (1993, 2001, 2008); Department of Zoology, University of British Columbia, Vancouver, Canada (1996); School of Biosciences, University of Birmingham, UK (2000); Department of Biological Sciences, University of Naples Federico II, Napoli, Italy (2005); School of Biological Sciences, University of Liverpool, UK (2008); Department of Biology, Turku University, Finland (2008); Department of Cell Biology, University of Calabria, Cosenza, Italy (2010). 

Referee work and external committees 
Referee for 40 different scientific journals and reviewer of grant proposals to Norwegian Research Council, Natural Environment Research Council UK, Biotechnology and Biological Sciences Research Council UK.
Member of several committees evaluating doctorial dissertations and/or applications for academic positions at universities in Denmark and abroad.
Member of the National Bibliometrics Research Indicator for Biology (January 2008 → December 2013).
Member of FNU, The Danish Natural Science Research Council (January 2011 → December 2014 ).

Teaching and supervision
Have taught courses on Physiological Adaptation to Environmental Stress, Ecological Physiology, General Physiology/Zoophysiology, Biology Experiments - Communicating Biology, Knowledge in Laboratory Animal Science, Biological Oceanography, Projects in Natural Science, Fish Ecophysiology, Acid Base Regulation, Biology of Oxygen Depletion, and 1-year course in Chemistry, Biology and Molecular Biology. 
Supervised 7 postdocs, 6 Ph.D. projects, 37 M.Sc. (cand. scient.) projects, 35 B.Sc. (bachelor) projects and 18 further experimental student projects. 

Administration (selected examples)
Member of various university study councils (1989-1992, 1994), member of the Ph.D. committee at Institute of Biology (Odense) 1991-2017, member of the Biology Institute Council 1993-1999, Institute leader 1996-1998.

Other selected activities
Organizer/co-organizer of nine international scientific meetings and workshops. Member of the Society for Experimental Biology, American Physiological Society, Scandinavian Physiological Society, European Society for Comparative Physiology and Biochemistry and Danish Academy of Natural Science.

Author/co-author of 117 peer-reviewed articles/chapters in international scientific journals and books, and editor of one book on “Fish Ecophysiology”. Over 100 abstracts from lectures and poster presentations given at various scientific meetings.

Research areas

Research areas

Comparative physiology, respiration, gas transport, acid-base balance, ionic regulation and cellular ion transport. The research aims at understanding the influence of environment on the physiology of animals, and it includes studies of mechanisms at molecular, cellular and organismic levels.

The research includes investigations on:
• Acute and chronic effects of environmental changes (oxygen shortage, hypercapnia, salinity change etc.) in fish and crustaceans.
• Functional properties of haemoglobins.
• Interaction of haemoglobin with membrane proteins.
• Blood gas transport.
• Ion transport, volume regulation and pH regulation in red blood cells.
• Circulatory and ventilatory responses to hypoxia.
• Comparative physiology of nitric oxide and its metabolites.
• The role of red blood cells in local blood flow regulation via oxygenation-dependent release of vasodilatory compounds (NO and ATP).
• The physiological role of nitrite as a signalling molecule and NO donor.
• Acid-base regulation in vertebrates and invertebrates.
• Physiological effects of contaminants (copper, ammonia, nitrite, nitrate).


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